This is an illustrated key to the families of Gymnosperms. These are woody plants, exclusively wind-pollinated, of very ancient origin, including many extinct families and genera.
The key feature is that the ovules are naked (left) not enclosed inside an ovary as in the Angiosperms (right). The ovules, and subsequently the seeds, are typically borne on the face of a cone scale. The pollen enters the micropyle directly, not via a pollen tube down the style
Palm-like or fern-like plants with a trunk less than 2m tall, composed of leaf bases supporting a rosette of large, compound leaves. These belong to the order Cycadales.
The order Cycadales comprises the families Cycadacea (e.g. the familiar Cycas revoluta; left), Zamiaceae (8 genera including Encephalartos; right) and Stangeriaceae (just Stangeria and Bowenia). Otherwise, next tweet.
Small shrubs of a switch-like habit are the order Gnetales, which contains some famous plants including Welwitschia (left; Welwitschiaceae), and Ephedra (right; Ephedraceae). Otherwise, next tweet.
Leaves broad (>2cm), fan-shaped, with a long petiole, deciduous (left). This is Ginkgoaceae, represented by the common urban street tree Ginkgo biloba. The buds on the winter twigs are highly distinctive (right).
Ginkgo trees are male (left) or female (right) i.e. dioecious. Males are typically grown because the fruits borne by females are unpleasantly smelly ("a nauseous stink" according to Stace). Plant not like this, next tweet.
Leaves stalked but narrowly linear lanceolate (i.e. not broad) up to 20cm x 3cm, with seeds typically inside a fleshy aril are Podocarpaceae. There are 6 genera. The commonest species are illustrated in this and the next 2 tweets. Dacrydium (left), Lagarostrobos (right).
Podocarpaceae genera II: Microcachrys (left), Phyllocladus (right)
Podocarpaceae genera III: Podocarpus (left, now known as Prumnopitys andina), and Saxegothea (right). Plants not like this, next tweet.
Leaves sessile, broad, many-veined and leathery, closely spiralling the stem, and buds lacking proper bud scales, are Araucariaceae.
The Monkey-puzzle Araucaria araucana (last tweet) is the only common tree outdoors in Britain, but the tree you remember from your Mediterranean holidays, A. heterophylla (right) survives the winter outside in London these days.
Shrubs with lanceolate leaves in 2 lateral ranks, tapering gradually to a sharp tip. This family is Cephalotaxacea. There are just two species, both in the genus Cephalotaxus: left is C. harringtonia (short leaves < 6cm); right is C. fortunei (longer leaves, up to 9cm).
If your plant is a bit more Yew-like, and has parallel-sided leaves with rounded, obtuse tips, and buds with green bud-scales, then look for fruits. If you find solitary seeds more or less enclosed in a red fleshy aril (i.e. a bit berry-like) then your plant is Taxaceae.
Our native is Taxus baccata, of which there are dozens of cultivars to be seen on any urban walk ('Fastigiata Aurea'; left)' 'Fastigiata'; right). There are 4 other species and 2 hybrids grown in British gardens, all of which are rare and well worth seeking out.
Finally, we come to the big two Gymnosperm families: Pinaceae and Cupressaceae. The Pinaceae have needles for leaves, either singly or in bundles. There are two sorts of Cupressaceae: redwoods (formerly Taxodiaceae) with awl-shaped leaves and cypresses with scale-like leaves.
These groups will get close attention both when we look at how to tell the genera apart (next thread), and then even more detail when we look at keys to the species within each of the genera. For now, just enjoy the families.
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This is an illustrated key to the genera of the Gymnosperm family Pinaceae. This is the biggest and in many ways the most important of the conifer families, and contains many of the really familiar kinds: pines, spruces, firs, larches, cedars and so forth.
The big groups within Pinaceae have needles in bundles (Pinus, Larix, Cedrus, Pseudolarix; left) or needles solitary (Picea, Abies, Pseudotsuga, Tsuga; right). You need to look closely enough to make absolutely sure that you have got this distinction right. Bundle or solitary ?
We’ll cover the needles in bundles first. If there are 5 or fewer needles per bundle, you have Pinus (illustrated). If more than this, then something else. Technically, the bundle is called a fascicle in Pinus, and is the membranous sheath at the base of these needle-groups.
This is an illustrated key to the genera of Gymnosperms. We have already covered several of them in the key to the families: Taxus (TL), Cephalotaxus (TR), Ginkgo (BL) and Araucaria (BR) with the various podocarps, cycads and gnetales, so we don’t need to do any more with these.
That just leaves 2 families to deal with. But what a pair of families they are: Pinaceae and Cupressaceae. The first thing to appreciate is just how variable the different genera within these families are from one another: here for instance, is Sequoiadendron (L) and Sequoia (R)
Again, still within Cupressaceae, here are the contrasting cones of Nootka Cypress (left, essentially spherical) and Eastern White-cedar (right; more cylindrical).
Bottom-of-the-Wall Flora revisited. This lockdown is starkly different for botanizing, because so little is in flower compared to the first lockdown last spring. Not to mention the darkness, which is always a bit of a damper on plant-hunting.
Still, this is a really interesting time for urban plants because nearly everything is visible, just as juveniles rather than adults. What’s more, seedlings are extremely fresh and perky-looking at this time of year.
Learning seedlings opens up a whole new world of botanical fun. But Identifying them isn’t as hard as you might first think, because there's a good chance you'll remember what species are here from having seen them in flower during the first lock-down.
Let’s start with Chenopodium by doing the easy ones first. These can be identified without any real risk of error by jizz alone. I’ll ignore the definitions of the 5 new genera for the time being.
Easiest of all is Strawberry-blite (Chenopodium capitatum) with its succulent, bright red tepals. This is by far the rarest of all the distinctive species.
Talking about colourful, the ornamental Quinoa are pretty unmistakable (Chenopodium quinoa). Green plants of this species, however, and much more tricky and certainly can’t be done by jizz.
Admit it. You’ve always intended to get to grips with Atriplex and Chenopodium, but somehow you just never got round to it. The time has come. The fruits are ripe and there are no more excuses. We’ll start with the species that you can do by jizz alone, in the field.
Starting with the easy Atriplex. There are two shrubby ones and two annual herbs. The native shrubby one is A. portulacoides. This is a a locally dominant sprawling plant of saline mud and sand, with lower leaves opposite. It is often flooded at high tide.
The alien shrubby one is Atriplex halimus. This is a tall alien shrub (up to 2.5m) planted as a seaside windbreak and occasionally naturalized. It has all of its leaves alternate.
Many finger-grasses were wool-aliens from Australia and South Africa. They were imported as seeds tangled up in the wool or embedded in the dags that clung to the tail and rear-end of the animal. Once importation of dirty fleeces stopped (mid 20th C) they were seldom recorded.
The digitate grass genera in each of the 4 categories are separated as follows. First the paired inflorescences. There are just two genera, both rare. Paspalum distichum (left) has glumes thinner than upper lemmas. Andropogon distachyos (right) has glumes thicker than lemmas.
Digitate genera: 1 common (Digitaria), 1 occasional (Cynodon), 3 rare (Eleusine, Chloris & Dactyloctenium). Telling Digitaria from Cynodon is tricky (x20 ideally). Count the scales beneath the fertile floret (glumes + lemmas + paleas). Cynodon = 2 (left); Digitaria = 3 or 4 (R).