1/ Overnight news of a devastating Ukrainian HIMARS strike against a Russian ammunition train suggests to me that the Ukrainians have been rather clever in exploiting the limitations of the local rail network.
2/ The attack took place at Brylivka railway station, south-east of Kherson. Coincidentally, it's an area I remember from a visit many years ago. The whole area is a vast, flat, arid and frankly mononous farming region watered by irrigation canals. 
3/ Brylivka owes its existence to the railway line, which was built in 1944 under Stalin to provide a second rail route to Crimea (the main line is further east, running from Melitopol to Simferopol). The village was founded the following year, presumably to house railway staff.
4/ But the line at Brylivka has three peculiarities. First and most importantly, the entire line from Kherson to Dzhankoy is only a single track line. Single track lines have a very limited capacity to carry trains. (Thanks to bueker.net for the map.)
5/ There had been a plan to upgrade the line to double tracks with electrification during the 2010s, but this fell through due to Russia's seizure of the Crimea in 2014.
6/ Second, Brylivka is equipped with a large set of passing loops (or passing sidings) which are long enough for large freight trains. Passing loops allow trains to pass in both directions on a single-line track. The Russian ammo train would have been stopped here.
7/ Third, Brylivka is just south of the North Crimean Canal, which waters the entire area (and Crimea). The railway line crosses it on a single-tack bridge – given its strategic importance, I wouldn't be surprised if Mr HIMARS paid it a visit soon.
8/ The line has not been very busy in recent years. Russia's takeover of the Crimea meant that long-distance and freight traffic ceased in 2014. Prior to the 2022 invasion, it reportedly only had 2 passenger trains a day between Kherson and Vadim, the last Ukrainian-held station.
9/ However, given Russia's dependency on railways for its military logistics (as noted by @TrentTelenko and others), the Russians are likely to have been making heavy use of the line to resupply their forces in occupied areas of Kherson oblast.
10/ They have also within the last month reopened the line from Kherson to Dzhankoy for passenger traffic, though I would imagine the timetable will be somewhat disrupted now. ria.ru/20220630/melit…
11/ So I think it's likely that the Ukrainians could predict where the ammo train would be stopping, because the single-track layout of the line likely required a stop at Brylivka's passing loops.
12/ The damage at Brylivka certainly looks severe, though I would imagine the Russians will be able to repair the track within a few days. But there's nothing they can do about the track's layout.
13/ Whatever else happens, the track is likely to remain single, there will continue to be a need for a passing loop at Brylivka, and trains will continue to need to stop there to allow other trains to pass. So this vulnerability isn't going to go away. /end
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