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It’s not at all clear that the Secretary of Defense even has the authority to fire the Secretary of the Navy—who’s appointed by the President under 10 U.S.C. § 8013(a)(1):

law.cornell.edu/uscode/text/10…

§ 8013 says he’s subject to _direction_ of SecDef, but says nothing about removal.
Of course, even if this is right, it’s a technicality—since (1) Spencer resigned anyway; and (2) the President unquestionably could fire him.

But once again, we get an object lesson in the technical but important law of appointing and removing executive branch officers.
And #SCOTUS has been clear since at least 1839 that, unless a statute specifically says otherwise, the appointing authority (here, the President) is the default removal authority:

caselaw.findlaw.com/us-supreme-cou…
And here's @realDonaldTrump's tweet in which he says Spencer's "services have been terminated" by _Esper_, something Esper had no legal authority to do (but Trump did):

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